2c78d69bf51c0282d97dd262b8f9005f1c5d3192
kate
  Tue May 12 16:32:36 2020 -0700
Polish track config and draft track description. refs #24668

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+<h2>Description</h2>
+<p>
+This track displays the <em>ENCODE Registry of candidate cis-Regulatory Elements</em> (cCREs) in
+the human genome, a total of 926,535 elements identified and classified by the 
+ENCODE Data Analysis Center according to regulatory effect.
+cCREs are the subset of DNase hypersensitivity sites clustered across all ENCODE samples
+that are supported by either histone modifications (H3K4me3 and H3K27ac) or CTCF-binding data.
+The cCRE dataset is the core of the integrative level of epigenomic and transcriptomic
+annotations produced by ENCODE.
+</p>
+Additional exploration of the cCRE's and underlying raw ENCODE data is provided by the
+<a target="_blank" href="https://screen.wenglab.org/">
+SCREEN</a>
+(Search Candidate cis-Regulatory Elements) web tool,
+designed specifically for the registry, accessible by linkouts from the track details page.
+
+<!--
+<p>
+The related cCREs by Biosample composite track presents ccREs 
+and associated epigenetic signal in all individual biosamples in a large matrix.
+Additional views of the data are provided by the <a>ENCODE Integrative Megahub</a>.
+</p>
+-->
+
+<h2>Display Conventions and Configuration</h2>
+<p>
+CCREs are colored according to classification by regulatory signature:
+<p>
+<table cellpadding='2'>
+  
+  <tr>
+    <th style="border-bottom: 2px solid;">Color</th>
+    <th style="border-bottom: 2px solid;"></th>
+    <th style="border-bottom: 2px solid;">Group</th>
+    <th style="border-bottom: 2px solid;"></th>
+  </tr>
+  </thead>
+
+<tr><td style='background-color: red;'></td><td>red</td>
+        <td>promoter-like</td>
+        <td>PLS</td></tr>
+<tr><td style='background-color: orange;'></td><td>orange</td>
+        <td>proximal enhancer-like</td>
+        <td>pELS</td></tr>
+<tr><td style='background-color: yellow;'></td><td>yellow</td>
+        <td>distal enhancer-like</td>
+        <td>dELS</td></tr>
+<tr><td style='background-color: blue;'</td><td>blue</td>
+        <td>CTCF-only</td>
+        <td>CTCF-only</td></tr>
+<tr><td style='background-color: #ffa0a0;'></td><td>pink</td>
+        <td>DNase-H3K4me3</td>
+        <td>DNase-H3K4me3</td></tr>
+</table>
+</p>
+
+<h2>Methods</h2>
+<p>
+All individual DNase hypsersensitivity sites (DHSs) identified from 706 DNAse-seq experiments
+in human (a total of 93 million) were iteratively clustered and filtered 
+for highest signal across all experiments, producing 2.2 million representative DHSs (rDHSs).
+The highest signal elements from this set that were also supported by high H3K4me3, H3K27ac 
+and/or CTCF ChIP-seq signals were designated cCRE's (a total of 926,535 in human).
+</p>
+<p>
+Classification of cCRE's was performed based on the following criteria:
+<p>
+<p>
+1. cCREs with promoter-like signatures (cCRE-PLS) fall within 200 bp of 
+an annotated GENCODE TSS and have high DNase and H3K4me3 signals.
+</p>
+<p>
+2. cCREs with enhancer-like signatures (cCRE-ELS) have high DNase and H3K27ac 
+with low H3K4me3 max-Z score if they are within 200 bp of an annotated TSS. 
+The subset of cCREs-ELS within 2 kb of a TSS is denoted proximal (cCRE-pELS), 
+while the remaining subset is denoted distal (cCRE-dELS).
+</p>
+
+<p>
+3. DNase-H3K4me3 cCREs have high H3K4me3 max-Z scores but low H3K27ac max-Z scores 
+and do not fall within 200 bp of a TSS.
+</p>
+
+<p>
+4. CTCF-only cCREs have high DNase and CTCF and low H3K4me3 and H3K27ac.
+</p>
+
+<img style='margin-left: 40px;' height=229 width=371 src="../images/cCREgroups.png">
+
+<p>
+For further detail about the identification and classification of ENCODE cCREs see 
+the <i>About</i> page of the
+<a target="_blank" href="https://screen.encodeproject.org">SCREEN</a> web tool.
+</p>
+
+<h2>Data Access</h2>
+<p>
+The ENCODE accession numbers of the constituent datasets at the
+<a target="_blank" href="https://encodeproject.org">ENCODE Portal</a>
+are available from the cCRE details page.
+</p>
+<p>
+The data in this track can be interactively explored with the 
+<a href="../cgi-bin/hgTables">Table Browser</a> or the 
+<a href="../cgi-bin/hgIntegrator">Data Integrator</a>. 
+The data can be accessed from scripts through our 
+<a href="https://api.genome.ucsc.edu">API</a>, the track name is "encodeCcreCombined".
+
+<p>
+For automated download and analysis, this annotation is stored in a bigBed file that
+can be downloaded from
+<a href="http://hgdownload.soe.ucsc.edu/gbdb/$db/bbi/" target="_blank">our download server</a>.
+The file for this track is called <tt>encodeCcreCombined.bb</tt>. 
+Individual regions or the whole genome annotation can be obtained using our tool 
+<tt>bigBedToBed</tt> which can be compiled from the source code or downloaded as a precompiled
+binary for your system. 
+Instructions for downloading source code and binaries can be found
+<a href="http://hgdownload.soe.ucsc.edu/downloads.html#utilities_downloads">here</a>.
+The tool can also be used to obtain only features within a given range, e.g. 
+<tt>bigBedToBed http://hgdownload.soe.ucsc.edu/gbdb/hg19/bbi/encodeCcreCombined.bb -chrom=chr21 -start=0 -end=100000000 stdout</tt></p>
+</p>
+
+<h2>Credits</h2>
+<p>
+This dataset was produced by the ENCODE Data Analysis Center (
+<a target="_blank" href="https://www.umassmed.edu/zlab/">Zlab</a>
+ at UMass Medical Center). 
+Thanks to Henry Pratt, Jill Moore, Michael Purcaro, and Zhiping Weng, PI for providing
+this data.
+Thanks also to the ENCODE Consortium, the ENCODE production laboratories, 
+and the ENCODE Data Coordination Center for generating and processing the datasets used here.
+</p>
+
+<h2>References</h2>
+<p>
+ENCODE Project Consortium..
+<a href="https://doi.org/10.1038/nature11247" target="_blank">
+An integrated encyclopedia of DNA elements in the human genome</a>.
+<em>Nature</em>. 2012 Sep 6;489(7414):57-74.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/22955616" target="_blank">22955616</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3439153/" target="_blank">PMC3439153</a>
+</p>
+
+ENCODE Project Consortium..
+<a href="http://dx.plos.org/10.1371/journal.pbio.1001046" target="_blank">
+A user's guide to the encyclopedia of DNA elements (ENCODE)</a>.
+<em>PLoS Biol</em>. 2011 Apr;9(4):e1001046.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/21526222" target="_blank">21526222</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3079585/" target="_blank">PMC3079585</a>
+</p>
+