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+<H2>Description</H2>
+<p>
+This track set shows the results of the
+<a target=_blank href="https://www.covid19hg.org/blog/2020-11-24-covid-19-hgi-results-for-data-freeze-4-october-2020/"<b>Data Release 4 (October 2020)</b></a>
+of <b>GWAS meta-analyses</b> from the 
+<a target=_blank href="https://www.covid19hg.org/">
+<b>COVID-19 Host Genetics Initiative (HGI)</b></a>: 
+a collaborative effort to facilitate 
+the generation, analysis and sharing of COVID-19 host genetics research.
+The COVID-19 HGI organizes meta-analyses across multiple studies contributed by 
+<a target="_blank" href="https://www.covid19hg.org/partners/">partners world-wide</a>
+to identify the genetic determinants of SARS-CoV-2 infection susceptibility and disease severity 
+and outcomes. Moreover, the COVID-19 HGI also aims to provide a platform for study partners to 
+share analytical results in the form of summary statistics and/or individual level data where 
+possible.
+At the time of this release, a total of 137 studies were registered with this effort.
+</p>
+
+<p>
+The specific phenotypes studied by the COVID-19 HGI are those that benefit from maximal sample 
+size: primary analysis on disease severity. For the Data Release 4 the number of cases have
+increased by nearly ten-fold (over 30,000 COVID-19 cases and 1.47 million controls) by combining
+data from 34 studies across 16 countries. The increased sample size resulted in strong evidence of 
+seven genomic regions associated with severe COVID-19, on chromosomes 3, 6, 9, 12, 19, and 21; and 
+one additional signal on chromosome 3 associated with COVID-19 partial-susceptibility.
+The four tracks here are based on data from HGI meta-analyses A2, B2, C1, and C2, described here:
+</p>
+
+<ul>
+<li>Severe COVID GWAS (<b>A2</b>): Very severe respiratory confirmed covid vs. population (4933 cases from 14 studies)</li>
+<li>Hosp COVID GWAS (<b>B2</b>): Hospitalized covid vs. population (8638 cases from 23 studies)</li>
+<li>Tested COVID GWAS (<b>C1</b>): Covid vs. lab/self-reported negative (8638 cases from 23 studies)</li>
+<li>COVID GWAS (<b>C2</b>): Covid vs. population (30937 cases from 36 studies)</li>
+</ul>
+
+Due to privacy concerns, these browser tracks exclude some of the data in the full analysis 
+results (specifically, data provided by 23andMe contributed studies). The actual study and case 
+and control counts for the individual browser tracks are listed in the track labels (shown 
+in the 'List subtracks' section above).
+
+
+<H2>Display Conventions</H2>
+<p>
+Displayed items are colored by <b>GWAS effect</b>:  red for positive (harmful) effect, 
+blue for negative (protective) effect.
+The height ('lollipop stem') of the item is based on statistical significance (<b>pvalue</b>) 
+or  effect size (<b>beta coefficient</b>). 
+For better visualization of the data, only SNPs with p-values smaller than 1e-3 are 
+displayed by default.</p>
+<p>
+For tracks based on effect size, the
+color saturation indicates statistical significance: p-values smaller than 1e-5
+are brightly colored (bright red
+<span style='background-color: #ff0000;'>&nbsp;&nbsp;</span>
+, bright blue
+<span style='background-color: #0000ff;'>&nbsp;&nbsp;</span>
+),
+those with less significance (p &gt;= 1e-5) are paler (light red
+<span style='background-color: #ffa0a0;'>&nbsp;&nbsp;</span>
+, light blue
+<span style='background-color: #a0a0ff;'>&nbsp;&nbsp;</span>
+). 
+For track based on pvalue, the color brightness reflects the effect size.</p>
+<p>
+Each track has separate display controls and data can be filtered according to the
+<b>number of studies</b>, <b>minimum -log10 p-value</b>, and the
+<b>effect size (beta coefficient)</b>, using the track <b>Configure</b> options.</p>
+<p>
+<b>Mouseover</b> on items shows the rs ID (or chrom:pos if none assigned), both the non-effect 
+and effect alleles, the effect size (beta coefficient), the p-value, and the number of 
+studies.
+Additional information on each variant can be found on the details page by clicking on 
+the item. </p>  
+
+<H2>Methods</H2>
+<p>
+COVID-19 Host Genetics Initiative (HGI) GWAS meta-analysis round 4 (October 2020) results were 
+used in this study. 
+Each participating study partner submitted GWAS summary statistics for up to four 
+of the <a target=_blank href="https://www.covid19hg.org/results/">COVID-19 phenotype definitions</a>.</p>
+<p>
+Data were generated from genome-wide SNP array and whole exome and genome
+sequencing, leveraging the impact of both common and rare variants. The statistical analysis
+performed takes into account differences between sex, ancestry, and date of sample collection. 
+Alleles were harmonized across studies and reported allele frequencies are based on gnomAD 
+version 3.0 reference data. Most study partners used the <b>SAIGE GWAS</b> pipeline in order 
+to generate summary statistics used for the COVID-19 HGI meta-analysis. The summary statistics 
+of individual studies were manually examined for inflation, 
+deflation, and excessive number of false positives.  
+Qualifying summary statistics were filtered for 
+<b>INFO > 0.6</b> and <b>MAF > 0.0001</b> prior to meta-analyzing the entirety of the data. 
+</p>
+The meta-analysis was performed using fixed effects inverse variance weighting.
+The meta-analysis software and workflow are available <a target=_blank 
+href="https://github.com/covid19-hg/META_ANALYSIS">here</a>. More information about the 
+prospective studies, processing pipeline, results and data sharing can be found 
+<a target=_blank href="https://www.covid19hg.org/about/">here</a>.
+</p>
+
+<H2>Data Access</H2>
+<p>
+The data underlying these tracks and summary statistics results are publicly available in <a target=_blank href="https://www.covid19hg.org/results">COVID19-hg Release 4 (October 2020)</a>.
+The raw data can be explored interactively with the <a target="_blank" href="../cgi-bin/hgTables">
+Table Browser</a>, or the <a target="_blank" href="../cgi-bin/hgIntegrator">Data Integrator</a>. 
+Please refer to
+our <a href="https://groups.google.com/a/soe.ucsc.edu/forum/#!forum/genome"
+target="_blank">mailing list archives</a> for questions, or our <a target="_blank"
+href="../FAQ/FAQdownloads.html#download36">Data Access FAQ</a> for more information.
+</p>
+
+<H2>Credits</H2>
+<p>
+Thanks to the COVID-19 Host Genetics Initiative contributors and project leads for making these 
+data available, and in particular to Rachel Liao, Juha Karjalainen, and Kumar Veerapen at the 
+Broad Institute for their review and input during browser track development.
+</p>
+
+<H2>References</H2>
+
+<p>
+COVID-19 Host Genetics Initiative.
+<a href="https://www.ncbi.nlm.nih.gov/pubmed/32404885" target="_blank">
+The COVID-19 Host Genetics Initiative, a global initiative to elucidate the role of host genetic
+factors in susceptibility and severity of the SARS-CoV-2 virus pandemic</a>.
+<em>Eur J Hum Genet</em>. 2020 Jun;28(6):715-718.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/32404885" target="_blank">32404885</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7220587/" target="_blank">PMC7220587</a>
+</p>
+
+
+