44c00f07b0e94306e09f30c84ea6ab0f044e1a29
max
  Fri Aug 14 05:09:12 2026 -0700
adding lin et al long-read SV subtrack, refs #38099

diff --git src/hg/makeDb/trackDb/human/lrSv1kLin.html src/hg/makeDb/trackDb/human/lrSv1kLin.html
index 05b20758df6..9d7706b33f3 100644
--- src/hg/makeDb/trackDb/human/lrSv1kLin.html
+++ src/hg/makeDb/trackDb/human/lrSv1kLin.html
@@ -1,114 +1,151 @@
 <h2>Description</h2>
 <p>
 This track shows structural variants (SVs) from an integrated long-read
 callset spanning 1,218 individuals of the 1000 Genomes Project. Structural
 variants are genomic rearrangements larger than about 50 bp, such as
 deletions and insertions; because they alter large stretches of DNA at once,
 they can affect gene dosage and regulation more strongly than
 single-nucleotide changes, and long reads resolve them far better than
 short-read data.
 </p>
 <p>
 Rather than coming from a single sequencing run, the calls are drawn together
-from several 1000 Genomes long-read efforts that use different technologies:
-HiFi and genome-assembly-based calls from the Human Pangenome Reference
-Consortium (<a href="hgTrackUi?g=hprc2v21Sv">HPRC year 2</a>), assembly-based
-calls from the <a href="hgTrackUi?g=hgsvc3Sv">Human Genome Structural
-Variation Consortium (HGSVC3)</a>, and Oxford Nanopore calls from the
-<a href="hgTrackUi?g=lrSv1kgOnt">Vienna 1000 Genomes ONT release</a>, together
-with Oxford Nanopore sequencing from the University of Washington 1000 Genomes
-ONT effort (see <a href="hgTrackUi?g=gustafsonSv">1KG ONT UW</a>). Sequencing
-of the 1000 Genomes collection is ongoing, so the number of individuals and
-variants in this track is expected to grow over time.
+from several 1000 Genomes long-read efforts that use different technologies.
+The 1,218 individuals combine:
 </p>
+<ul>
+<li>293 near-T2T haplotype-resolved assemblies from the Human Pangenome
+Reference Consortium (<a href="hgTrackUi?g=hprc2v21Sv">HPRC year 2</a>) and the
+<a href="hgTrackUi?g=hgsvc3Sv">Human Genome Structural Variation Consortium
+(HGSVC)</a>;</li>
+<li>480 roughly 30x Oxford Nanopore genomes generated by the 1000 Genomes
+Long Read Sequencing Consortium at the University of Washington (383 newly
+sequenced for this study and 97 previously published by
+<a href="hgTrackUi?g=gustafsonSv">Gustafson et al.</a>);</li>
+<li>445 low-pass Oxford Nanopore genomes retained after quality control from
+the <a href="hgTrackUi?g=lrSv1kgOnt">Vienna 1000 Genomes ONT release</a>
+(Schloissnig et al.).</li>
+</ul>
 <p>
-This track is preliminary and unpublished; its sample composition and variant
-counts will be updated as more long-read data becomes available.
-</p>
-<p>
-The current release contains more than 580,000 SVs on GRCh38 (about 391,000
-insertions and 196,000 deletions), each annotated with an overall allele
-frequency and allele frequencies for the five 1000 Genomes superpopulations
-(African, Admixed American, East Asian, European, South Asian). This is a
-preliminary, unpublished callset; the counts and sample composition will be
-updated as more data is added.
+The current release contains more than 580,000 SVs on GRCh38 (391,410
+insertions and 196,369 deletions) and more than 610,000 SVs on T2T-CHM13
+(376,117 insertions and 238,405 deletions), each annotated with an overall
+allele frequency and allele frequencies for the five 1000 Genomes
+superpopulations (African, Admixed American, East Asian, European, South
+Asian).
 </p>
 
 <h2>Display Conventions and Configuration</h2>
 <p>
 Items are colored by SV type, matching the other subtracks of the container:
 </p>
 <table class="stdTbl">
   <tr><th style="background-color:#C80000;width:2em">&nbsp;</th>
       <td>Deletion (DEL)</td></tr>
   <tr><th style="background-color:#0000C8;width:2em">&nbsp;</th>
       <td>Insertion (INS)</td></tr>
 </table>
 <p>
 Insertions are drawn at the insertion site with a width of 1 bp, and the
 length of inserted sequence is reported as the insertion length; deletions
 span the affected reference interval. The mouseover shows the variant name,
 SV type, reference and insertion lengths, allele count and per-population
 allele frequencies. Filters are available for SV type, SV length, insertion
 length, allele count, and overall and per-population allele frequency.
 </p>
 
 <h2>Methods</h2>
 <p>
-Per-sample long-read SV calls from the contributing 1000 Genomes efforts
-(HiFi and assembly-based calls from HPRC year 2 and HGSVC3, and Oxford
-Nanopore calls from the Vienna and University of Washington releases) were
-combined across the 1,218 individuals and merged into a single site-level
-callset with Truvari v5.2.0. Overall and per-superpopulation allele
-frequencies (EUR, AMR, EAS, AFR, SAS) were then added with bcftools
-fill-tags. Only deletions and insertions are reported in the current release.
-The callset is provided on both GRCh38/hg38 and T2T-CHM13/hs1 from the
-respective native assemblies.
+Lin et al. built the callset in two tiers. A baseline set of euchromatic SVs
+was established from 293 near-T2T haplotype-resolved assemblies generated by
+HGSVC and HPRC, and this was expanded with an additional 445 low-pass Oxford
+Nanopore genomes (Schloissnig et al. 2025) and 480 roughly 30x Oxford Nanopore
+genomes sequenced by the 1000 Genomes Long Read Sequencing Consortium (383 new
+plus 97 from Gustafson et al. 2024), for 1,218 genomes of diverse ancestry.
+Structural variants were discovered with ten long-read callers, and a
+machine-learning tool, BoostSV, ranked and selected the best allele to
+represent each SV across the different platforms and coverages, producing a
+single nonredundant callset. The final callset contains 614,522 SVs (376,117
+insertions and 238,405 deletions) on T2T-CHM13 and 587,779 SVs (391,410
+insertions and 196,369 deletions) on GRCh38. See Lin et al. for full details.
 </p>
 <p>
-The data was provided by the laboratories of Evan Eichler and Danny Miller
-(University of Washington) and is preliminary and unpublished; a manuscript
-is in preparation. The step-by-step build commands (format conversion and
-bigBed build) are recorded in the UCSC makeDoc for this track container:
+The insertion/deletion callset VCFs (GRCh38 and T2T-CHM13 native), already
+annotated with overall and per-superpopulation allele frequencies (EUR, AMR,
+EAS, AFR, SAS), were provided by the laboratories of Evan Eichler and Danny
+Miller (University of Washington). At UCSC the deletion and insertion records
+were converted to bigBed; no re-merging or re-annotation was performed. The
+step-by-step build commands (format conversion and bigBed build) are recorded
+in the UCSC makeDoc for this track container:
 <a href="https://github.com/ucscGenomeBrowser/kent/blob/master/src/hg/makeDb/doc/hg38/lrSv.txt" target="_blank">
 doc/hg38/lrSv.txt</a>. The conversion script and autoSql schema live in
 <a href="https://github.com/ucscGenomeBrowser/kent/tree/master/src/hg/makeDb/scripts/lrSv" target="_blank">
 makeDb/scripts/lrSv</a>, and the track configuration is in
 <a href="https://github.com/ucscGenomeBrowser/kent/blob/master/src/hg/makeDb/trackDb/human/lrSv.ra" target="_blank">trackDb/human/lrSv.ra</a>.
 </p>
 
 <h2>Data Access</h2>
 <p>
 The data can be explored interactively in table format with the
 <a href="../cgi-bin/hgTables">Table Browser</a> or the
 <a href="../cgi-bin/hgIntegrator">Data Integrator</a> and exported from there
 to spreadsheet or tab-separated tables. From scripts, the data can be accessed
 through our <a href="https://api.genome.ucsc.edu">API</a>, track=<i>lrSv1kLin</i>.
 </p>
 <p>
 For automated download and analysis, the annotation is stored in bigBed files
 that can be downloaded from our download server:
 <a href="http://hgdownload.soe.ucsc.edu/gbdb/hg38/lrSv/lin1218.bb" target="_blank">
 http://hgdownload.soe.ucsc.edu/gbdb/hg38/lrSv/lin1218.bb</a> (GRCh38/hg38,
 native) and
 <a href="http://hgdownload.soe.ucsc.edu/gbdb/hs1/lrSv/lin1218.bb" target="_blank">
 http://hgdownload.soe.ucsc.edu/gbdb/hs1/lrSv/lin1218.bb</a> (T2T-CHM13/hs1,
 native). Individual regions or the whole annotation can be obtained with the
 <tt>bigBedToBed</tt> utility, which can be compiled from source or downloaded
 as a precompiled binary from our
 <a href="http://hgdownload.soe.ucsc.edu/downloads.html#utilities_downloads" target="_blank">utilities
 page</a>. The tool can also extract features within a given range, for example:
 <tt>bigBedToBed http://hgdownload.soe.ucsc.edu/gbdb/hg38/lrSv/lin1218.bb -chrom=chr21 -start=0 -end=100000000 stdout</tt>.
 </p>
+<h2>Credits</h2>
 <p>
-This is a preliminary, unpublished callset provided by the authors; the
-underlying per-sample sequencing data is not yet publicly released.
+Thanks to Jiadong Lin for providing the merged callset and the dataset
+overview, and to Evan Eichler, Danny Miller and colleagues at the University of
+Washington, together with the contributing 1000 Genomes long-read consortia
+(HPRC, HGSVC and the 1000 Genomes ONT sequencing groups), for generating and
+sharing this callset.
 </p>
 
-<h2>Credits</h2>
+<h2>References</h2>
+<p>
+Lin J, <em>et al</em>. A high-resolution human pangenome structural variant
+resource for improved disease association. Submitted.
+</p>
+<p>
+Schloissnig S, Pani S, Ebler J, Hain C, Tsapalou V, S&#246;ylev A, H&#252;ther P, Ashraf H, Prodanov T,
+Asparuhova M <em>et al</em>.
+<a href="https://doi.org/10.1038/s41586-025-09290-7" target="_blank">
+Structural variation in 1,019 diverse humans based on long-read sequencing</a>.
+<em>Nature</em>. 2025 Aug;644(8076):442-452.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/40702182" target="_blank">40702182</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC12350158/" target="_blank">PMC12350158</a>
+</p>
+<p>
+Gustafson JA, Gibson SB, Damaraju N, Zalusky MPG, Hoekzema K, Twesigomwe D, Yang L, Snead AA,
+Richmond PA, De Coster W <em>et al</em>.
+<a href="http://genome.cshlp.org/lookup/pmidlookup?view=long&amp;pmid=39358015" target="_blank">
+High-coverage nanopore sequencing of samples from the 1000 Genomes Project to build a comprehensive
+catalog of human genetic variation</a>.
+<em>Genome Res</em>. 2024 Nov 20;34(11):2061-2073.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/39358015" target="_blank">39358015</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC11610458/" target="_blank">PMC11610458</a>
+</p>
 <p>
-Thanks to Evan Eichler, Danny Miller and colleagues at the University of
-Washington, and to the contributing 1000 Genomes long-read consortia (HPRC,
-HGSVC and the 1000 Genomes ONT sequencing groups), for generating and sharing
-this callset.
+Logsdon GA, Ebert P, Audano PA, Loftus M, Porubsky D, Ebler J, Yilmaz F, Hallast P, Prodanov T, Yoo
+D <em>et al</em>.
+<a href="https://doi.org/10.1038/s41586-025-09140-6" target="_blank">
+Complex genetic variation in nearly complete human genomes</a>.
+<em>Nature</em>. 2025 Aug;644(8076):430-441.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/40702183" target="_blank">40702183</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC12350169/" target="_blank">PMC12350169</a>
 </p>