68c5b3b5dfc4053ff78a6b1d236bd1ac90251cfa
lrnassar
  Mon Jun 1 14:40:45 2026 -0700
varFreqs: description pages for the three combined tracks and "SNV" rename
sweep.

Add varFreqsDisease.html and varFreqsArray.html so the two new combined
tracks have full Description/Display/Methods/Data Access/References. Add a
Caveats section on varFreqsArray about chip-data quality vs sequencing.

Update varFreqsAll.html and the supertrack varFreqs.html to reflect the
three-combined-track family (cross-links between siblings, new "Combined
Tracks" section, new table rows, and updated source/variant counts). Add a
GoNL row to the supertrack table.

Sweep 37 subtrack longLabels and four cross-referencing description pages
(colorsDbSnv.html, mei.html, meiSwegen.html, phasedVars.html) from
"Variant Frequencies:" to "SNV Frequencies:" to match the supertrack
shortLabel. refs #36642

diff --git src/hg/makeDb/trackDb/human/varFreqsDisease.html src/hg/makeDb/trackDb/human/varFreqsDisease.html
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+<h2>Description</h2>
+<p>
+This track merges variants from six disease-focused or clinically-recruited cohorts into a
+single bigBed file with predicted protein consequences and cross-database filtering. It
+contains 932 million variants from SFARI SPARK (WES + WGS, autism families), TOPMed
+(NHLBI heart, lung and blood disease cohorts), SCHEMA (schizophrenia case/control),
+GREGoR (rare-disease families), and GA4K (PacBio long-read pediatric rare disease). Where
+the source dataset provides per-phenotype counts, those are exposed as separate AC/AF
+columns and as filter widgets.
+</p>
+
+<p>
+For a summary of all available variant frequency databases, including the population-scale
+control track and the genotyping-array track, see the
+<a href="hgTrackUi?g=varFreqs">SNV Frequencies</a> supertrack page.
+</p>
+
+<p>
+Each variant is annotated with its predicted consequence on protein-coding genes
+(using <a href="https://samtools.github.io/bcftools/howtos/csq-calling.html"
+target="_blank">bcftools csq</a> with
+<a href="https://www.ensembl.org/info/data/ftp/index.html" target="_blank">Ensembl</a>
+gene models), and colored by severity. Allele counts and frequencies are shown for each
+source database and, where available, broken down by phenotype.
+</p>
+
+<h2>Display Conventions</h2>
+
+<h3>Color by Consequence</h3>
+<p>Variants are colored by their most severe predicted consequence:</p>
+<table class="stdTbl">
+<tr><th>Color</th><th>Consequence class</th><th>Examples</th></tr>
+<tr>
+  <td style="background-color: rgb(255,0,0); color: white; text-align: center;"><b>Red</b></td>
+  <td>Protein-truncating / Loss-of-function</td>
+  <td>stop_gained, frameshift, splice_donor, splice_acceptor, stop_lost, start_lost</td>
+</tr>
+<tr>
+  <td style="background-color: rgb(31,119,180); color: white; text-align: center;"><b>Blue</b></td>
+  <td>Missense / In-frame</td>
+  <td>missense, inframe_insertion, inframe_deletion, protein_altering</td>
+</tr>
+<tr>
+  <td style="background-color: rgb(0,128,0); color: white; text-align: center;"><b>Green</b></td>
+  <td>Synonymous</td>
+  <td>synonymous, stop_retained</td>
+</tr>
+<tr>
+  <td style="background-color: rgb(128,128,128); color: white; text-align: center;"><b>Grey</b></td>
+  <td>Non-coding / Intergenic</td>
+  <td>intron, non_coding, intergenic, UTR</td>
+</tr>
+</table>
+
+<h3>Amino Acid Change Notation</h3>
+<p>
+The &quot;AA change&quot; field uses bcftools csq notation: <b>23I&gt;23V</b> means position
+23 changed from Isoleucine (I) to Valine (V) (missense). <b>23I</b> alone (no arrow)
+means position 23 is Isoleucine and unchanged (synonymous). A &quot;*&quot; indicates a
+stop codon (e.g. 45R&gt;45* is a stop_gained).
+</p>
+
+<h2>Filters</h2>
+<p>
+This track supports filtering via the track settings page. Click the track title or use the
+&quot;Configure&quot; button to access filters.
+</p>
+
+<h3>Variant Type and Consequence</h3>
+<ul>
+  <li><b>Variant Type</b>: SNV, Insertion, Deletion, or MNV.</li>
+  <li><b>Consequence</b>: Missense, Synonymous, Stop Gained, Frameshift, Splice Donor,
+      Splice Acceptor, Intron, 3' UTR, 5' UTR, Non-coding, Intergenic, or Other. The filter
+      uses OR logic across the comma-separated consequence tokens on each variant. See the
+      <a href="hgTrackUi?g=varFreqsAll">All Databases Combined</a> description page for a
+      complete description of the &quot;Other&quot; bucket.</li>
+</ul>
+
+<h3>Frequency and Count Filters</h3>
+<ul>
+  <li><b>Max Allele Frequency</b>: Filter by the maximum allele frequency observed across
+      the six disease cohorts. Useful for finding rare variants enriched in cases.</li>
+  <li><b>Total Allele Count</b>: Filter by the sum of allele counts across all six
+      databases.</li>
+  <li><b>Per-database AF and AC</b>: Filter by allele frequency or count in any specific
+      source. For example, restrict to variants with SCHEMA case AF &gt; 0.001.</li>
+</ul>
+
+<h3>Phenotype-stratified Filters</h3>
+<p>
+Four of the six sources publish counts split by phenotype, which lets you compare allele
+frequencies between affected and unaffected groups within the same cohort:
+</p>
+<ul>
+  <li><b>SPARK WES</b> and <b>SFARI WGS</b>: ASD proband counts versus non-ASD family
+      members (mostly parents and unaffected siblings). The split is from the SPARK
+      individuals_registration <code>asd</code> column.</li>
+  <li><b>SCHEMA</b>: Schizophrenia case counts versus controls, summed across the 39
+      analysis cohorts in the original meta-analysis.</li>
+  <li><b>GREGoR</b>: Affected, Unaffected, and Unknown disease-status counts.</li>
+</ul>
+
+<h3>Source Database</h3>
+<p>
+The <b>Source Database</b> filter restricts the display to variants present in specific
+databases. It uses OR logic: selecting multiple databases shows variants found in any of
+the selected sources.
+</p>
+
+<h3>Length Filters</h3>
+<ul>
+  <li><b>Reference/Alternate Length</b>: Filter by the length of the reference or alternate allele.</li>
+  <li><b>Length Change</b>: Filter by the size difference between alternate and reference
+      (positive = insertion, negative = deletion, zero = SNV or MNV).</li>
+</ul>
+
+<h2>Methods</h2>
+<p>
+The same merge-and-annotate pipeline used for the
+<a href="hgTrackUi?g=varFreqsAll">All Databases Combined</a> track was run on the
+disease-cohort subset of source VCFs. Each VCF was stripped of its INFO fields,
+normalized with <code>bcftools norm</code> (splitting multi-allelic sites), and merged with
+<code>bcftools merge</code>. The merged VCF was then annotated with predicted protein
+consequences using <code>bcftools csq</code> with the
+<a href="https://www.ensembl.org/info/data/ftp/index.html" target="_blank">Ensembl</a>
+GRCh38 release 115 gene annotation (GFF3).
+</p>
+
+<p>
+The SPARK WES and WGS sites VCFs were rebuilt for this track so each variant carries
+phenotype-stratified counts in addition to overall AC/AN/AF. The split uses the
+<code>asd</code> column of the SPARK <code>individuals_registration</code> TSV via
+<code>bcftools +fill-tags -S</code>, producing AC_AUT / AN_AUT / AF_AUT and
+AC_NON_AUT / AN_NON_AUT / AF_NON_AUT. SCHEMA was processed the same way, summing
+AC_CASE/AN_CASE/AF_CASE and AC_CTRL/AN_CTRL/AF_CTRL across its 39 analysis cohorts.
+GREGoR ships AC/AN/AF triples for affected, unaffected and unknown disease status
+directly in its release.
+</p>
+
+<p>
+The track's
+<a href="https://github.com/ucscGenomeBrowser/kent/blob/master/src/hg/makeDb/doc/hg38/varFreqs.txt"
+target="_blank">makeDoc file</a> documents how each source VCF was converted. Scripts are
+available from
+<a href="https://github.com/ucscGenomeBrowser/kent/tree/master/src/hg/makeDb/scripts/varFreqs"
+target="_blank">Github</a>.
+</p>
+
+<h2>Data Access</h2>
+<p>
+The data can be explored interactively with the
+<a href="../cgi-bin/hgTables">Table Browser</a> or the
+<a href="../cgi-bin/hgIntegrator">Data Integrator</a>. For programmatic access, our
+<a href="https://api.genome.ucsc.edu" target="_blank">REST API</a> can be used; the track
+name is <em>varFreqsDisease</em>.
+</p>
+<p>
+Because the merged callset includes data from multiple sources whose redistribution
+licenses differ, the combined bigBed is <b>not available for download</b> from our download
+server. The combined track can be reconstructed from the individual source VCFs using the
+<a href="https://github.com/ucscGenomeBrowser/kent/tree/master/src/hg/makeDb/scripts/varFreqs"
+target="_blank">conversion scripts on GitHub</a> together with the
+<a href="https://github.com/ucscGenomeBrowser/kent/blob/master/src/hg/makeDb/doc/hg38/varFreqs.txt"
+target="_blank">build documentation</a>.
+</p>
+
+<h2>Credits</h2>
+<p>
+This track is only possible thanks to the data from millions of volunteers around the
+world, who donated blood, signed consent forms and provided health information about
+themselves and sometimes their families. Click on any of the individual tracks in the
+<a href="hgTrackUi?g=varFreqs">SNV Frequencies</a> supertrack to see the specific credits
+for each project. Thanks to Alex Ioannidis, UCSC, for the motivation for this track and
+to Andreas Lahner, MGZ, for feedback.
+</p>
+
+<h2>References</h2>
+<p>
+For primary citations of each source dataset, see the References section on the
+<a href="hgTrackUi?g=varFreqs">SNV Frequencies</a> supertrack page. The merged-track
+build itself uses the following tools:
+</p>
+<p>
+Danecek P, McCarthy SA.
+<a href="https://doi.org/10.1093/bioinformatics/btx100" target="_blank">
+BCFtools/csq: haplotype-aware variant consequences</a>.
+<em>Bioinformatics</em>. 2017 Jul 1;33(13):2037-2039.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/28205675" target="_blank">28205675</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5870570/" target="_blank">PMC5870570</a>
+</p>
+<p>
+McLaren W, Gil L, Hunt SE, Riat HS, Ritchie GR, Thormann A, Flicek P, Cunningham F.
+<a href="https://doi.org/10.1186/s13059-016-0974-4" target="_blank">
+The Ensembl Variant Effect Predictor</a>.
+<em>Genome Biol</em>. 2016 Jun 6;17(1):122.
+PMID: <a href="https://www.ncbi.nlm.nih.gov/pubmed/27268795" target="_blank">27268795</a>; PMC: <a
+href="https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4893825/" target="_blank">PMC4893825</a>
+</p>