691a2b8981d6db69e8707ea44041c4661cdac97e
max
Wed Sep 9 06:38:29 2026 -0700
Imprinting: add the ASM Atlas tracks, and tidy the collection's labels
Adds a composite built from Rosenski et al. 2025, "Atlas of imprinted and
allele-specific DNA methylation in the human body". Three subtracks: the
458 regions whose methylation follows the parent of origin, the 72 known
control regions with the boundaries the paper redrew, and the pool of
385,235 regions carrying two methylation states that those came out of.
A fourth set, the regions whose methylation follows a nearby SNP, is
built by the scripts but its stanza is commented out, since sequence
driven methylation is not imprinting.
The authors released hg19 only, so all three are lifted. Their published
files are close to bare BED, so the SNPs, cell types, p-values, gene
links and gamete methylation on the details pages are read out of the
paper's supplementary tables and joined on by position. Regions that
lift but change length by more than 10%, because hg38 added sequence
inside them, are kept with a note rather than dropped: one of them is
TCEB3C, the only control region on chr18.
Also across the collection:
- long labels name their source right after "Imprinting", so that a
label read on its own says where the data came from
- the two gene catalogs are worded alike, and ordered OMIM, Geneimprint,
MethBase2, Akbari, ASM Atlas
- the OMIM curators confirmed that their (I) marker covers established
and candidate imprinted genes alike, with nothing in the export to
tell them apart. Labels, description page and makeDoc now say so, and
the claim that the set is "more conservative" than the computational
tracks is gone. The bigBed was rebuilt for the autoSql line, same 459
features.
- every subtrack page opens by naming the collection, linked back to
its hgTrackUi page, and no longer repeats the collection page's
introduction to imprinting
refs #37599
diff --git src/hg/makeDb/trackDb/human/hg38/imprinting.ra src/hg/makeDb/trackDb/human/hg38/imprinting.ra
index 94a70e856f0..37970bfb851 100644
--- src/hg/makeDb/trackDb/human/hg38/imprinting.ra
+++ src/hg/makeDb/trackDb/human/hg38/imprinting.ra
@@ -1,90 +1,189 @@
track imprinting
shortLabel Imprinting
longLabel Genomic imprinting: allele-specific methylation and imprinted regions
group regulation
superTrack on
priority 3.5
track methBaseAsm
parent imprinting on
shortLabel MethBase2 ASM
- longLabel Imprinting: fraction of MethBase2 methylomes with allele-specific methylation at each CpG
+ longLabel Imprinting - MethBase2: fraction of methylomes with allele-specific methylation at each CpG
type bigWig 0 1
bigDataUrl /gbdb/hg38/imprinting/methBaseAsm/hg38.asm.bw
visibility full
- priority 1
+ priority 3
color 85,85,85
autoScale off
viewLimits 0:0.5
viewLimitsMax 0:1
maxHeightPixels 100:40:8
alwaysZero on
yLineOnOff on
yLineMark 0
windowingFunction mean
track geneimprint
parent imprinting on
- shortLabel Geneimprint
- longLabel Imprinting: Geneimprint catalog of imprinted and candidate imprinted genes
+ shortLabel Geneimprint Genes
+ longLabel Imprinting - Geneimprint: genes curated as imprinted or candidate imprinted
type bigBed 9 +
bigDataUrl /gbdb/hg38/imprinting/geneimprint/geneimprint.bb
visibility pack
priority 2
itemRgb on
searchIndex name
labelFields name,hgncSymbol
defaultLabelFields name
url https://geneimprint.com/site/genes/Homo_sapiens_$$
urlLabel Geneimprint entry for this gene:
urls hgncId="https://www.genenames.org/data/gene-symbol-report/#!/hgnc_id/$$"
mouseOver ${name}
Imprint status: ${status}
Expressed allele: ${expressedAllele}
Band in catalog: ${catalogBand}
Resolved to: ${hgncSymbol} (${resolvedBy})
filterValues.status Imprinted,Predicted,Not Imprinted,Conflicting Data,Provisional Data,Tissue Dependent,Unknown
filterType.status multipleListOr
filterLabel.status Filter by imprint status
filterValues.expressedAllele Paternal,Maternal,Biallelic,Isoform Dependent,Random,Unknown
filterType.expressedAllele multipleListOr
filterLabel.expressedAllele Filter by expressed allele
track akbariIdmr
parent imprinting on
shortLabel Akbari iDMRs
- longLabel Imprinting: imprinted differentially methylated regions compiled from five genome-wide studies
+ longLabel Imprinting - Akbari et al.: imprinted DMRs compiled from five genome-wide studies
type bigBed 9 +
bigDataUrl /gbdb/hg38/imprinting/akbari2023/akbariIdmr.bb
visibility pack
- priority 3
+ priority 4
itemRgb on
searchIndex name,gene
labelFields gene,name
defaultLabelFields gene
skipEmptyFields on
mouseOver ${name}
Methylated allele: ${methylatedAllele}
Evidence: ${evidence}
Reported by: ${studies}
filterValues.methylatedAllele Maternal,Paternal
filterType.methylatedAllele multipleListOr
filterLabel.methylatedAllele Filter by methylated allele
filterValues.evidence Multiple studies,Single study
filterType.evidence multipleListOr
filterLabel.evidence Filter by evidence
filterValues.studies Akbari,Court,Hernandez,Joshi,Zink
filterType.studies multipleListOr
filterLabel.studies Filter by source study
track omimImprint
parent imprinting on
- shortLabel OMIM Imprinted
- longLabel Imprinting: genes curated as imprinted by OMIM, from GeneScout
+ shortLabel OMIM Genes
+ longLabel Imprinting - OMIM: genes curated as imprinted or candidate imprinted, from GeneScout
type bigBed 9 +
bigDataUrl /gbdb/hg38/imprinting/omimImprint/omimImprint.bb
visibility pack
- priority 4
+ priority 1
itemRgb on
searchIndex name
url https://omim.org/entry/$$
urlLabel OMIM entry:
urls mimNumber="https://omim.org/entry/$$"
skipEmptyFields on
mouseOver ${name}
${geneName}
OMIM: ${entryType}
Band: ${cytoLocation}
filterValues.entryType OMIM gene,No OMIM entry
filterType.entryType multipleListOr
filterLabel.entryType Filter by whether OMIM has a gene entry
+
+ track kaplanImprint
+ parent imprinting on
+ compositeTrack on
+ shortLabel ASM Atlas
+ longLabel Imprinting - ASM Atlas: allele-specific methylation in 39 cell types (Rosenski 2025)
+ type bigBed 9 +
+ visibility hide
+ priority 5
+ itemRgb on
+ skipEmptyFields on
+ pennantIcon 19.jpg ../goldenPath/help/liftOver.html "lifted from hg19"
+
+ track kaplanParentalAsm
+ parent kaplanImprint on
+ shortLabel Parental ASM
+ longLabel Imprinting - ASM Atlas: regions where the methylation state follows the parent of origin
+ type bigBed 9 +
+ bigDataUrl /gbdb/hg38/imprinting/kaplanImprint/kaplanParentalAsm.bb
+ visibility pack
+ priority 1
+ searchIndex name
+ labelFields name,icrName
+ defaultLabelFields name
+ urls snps="https://www.ncbi.nlm.nih.gov/snp/$$"
+ mouseOver ${name}
Region: ${regionType}
Type: ${icrType}
Imprinted genes nearby: ${imprintedGenes}
Cell types: ${cellTypeCount}
Methylation in oocyte / sperm: ${methOocyte} / ${methSperm}
+ filterValues.regionType Known imprinting control region,Novel region near an imprinted gene,Novel region
+ filterType.regionType multipleListOr
+ filterLabel.regionType Filter by how the region relates to known imprinting
+ filterValues.icrType Oocyte gDMR,Sperm gDMR,Secondary DMR,Oocyte gDMR- secondary DMR,Sperm gDMR-secondary DMR,unclear
+ filterType.icrType multipleListOr
+ filterLabel.icrType Filter by the gamete the methylation comes from
+ filterByRange.cellTypeCount on
+ filter.cellTypeCount 1:73
+ filterLimits.cellTypeCount 1:73
+ filterLabel.cellTypeCount Number of cell types supporting the region
+ filterByRange.snpCount on
+ filter.snpCount 1:39
+ filterLimits.snpCount 1:39
+ filterLabel.snpCount Number of SNPs used to phase the region
+
+ track kaplanIcr
+ parent kaplanImprint on
+ shortLabel ICRs Revised
+ longLabel Imprinting - ASM Atlas: control regions with boundaries revised from fragment-level methylation
+ type bigBed 9 +
+ bigDataUrl /gbdb/hg38/imprinting/kaplanImprint/kaplanIcr.bb
+ visibility pack
+ priority 2
+ searchIndex name
+ labelFields name
+ defaultLabelFields name
+ mouseOver ${name}
Type: ${icrType}
Associated genes: ${genes}
Recovered as parent-of-origin ASM: ${parentalAsm}
+ filterValues.icrType Oocyte gDMR,Sperm gDMR,Secondary DMR,Oocyte gDMR- secondary DMR
+ filterType.icrType multipleListOr
+ filterLabel.icrType Filter by the gamete the methylation comes from
+ filterValues.parentalAsm yes,no
+ filterType.parentalAsm multipleListOr
+ filterLabel.parentalAsm Filter by whether this study recovered the region
+
+# track kaplanAsm
+# parent kaplanImprint off
+# shortLabel SNP-linked ASM
+# longLabel Imprinting - ASM Atlas: bimodal regions where methylation follows a heterozygous SNP
+# type bigBed 9 +
+# bigDataUrl /gbdb/hg38/imprinting/kaplanImprint/kaplanAsm.bb
+# visibility dense
+# priority 3
+# labelFields name
+# defaultLabelFields name
+# urls snps="https://www.ncbi.nlm.nih.gov/snp/$$"
+# mouseOver ${name}
ASM SNPs: ${snps}
Alleles: ${alleles}
Cell types: ${cellTypeCount}
Best adjusted p: ${minAdjP}
+# filterByRange.snpCount on
+# filter.snpCount 1:63
+# filterLimits.snpCount 1:63
+# filterLabel.snpCount Number of ASM SNPs in the region
+# filterByRange.cellTypeCount on
+# filter.cellTypeCount 1:81
+# filterLimits.cellTypeCount 1:81
+# filterLabel.cellTypeCount Number of cell types supporting the region
+
+ track kaplanBimodal
+ parent kaplanImprint off
+ shortLabel Bimodal Meth
+ longLabel Imprinting - ASM Atlas: regions where DNA fragments carry two distinct methylation states
+ type bigBed 9 +
+ bigDataUrl /gbdb/hg38/imprinting/kaplanImprint/kaplanBimodal.bb
+ visibility dense
+ priority 4
+ labelFields name
+ defaultLabelFields name
+ mouseOver ${name}
${cellTypes}
+ filterValues.cellTypes Adipocytes,Bladder-Ep,Blood-B,Blood-Granul,Blood-Mono+Macro,Blood-NK,Blood-T,Bone-Osteob,Breast-Basal-Ep,Breast-Luminal-Ep,Colon-Ep,Colon-Fibro,Dermal-Fibro,Endothel,Epid-Kerat,Eryth-prog,Fallopian-Ep,Gallbladder,Gastric-Ep,Head-Neck-Ep,Heart-Cardio,Heart-Fibro,Kidney-Ep,Liver-Hep,Lung-Ep-Alveo,Lung-Ep-Bron,Neuron,Oligodend,Ovary-Ep,Pancreas-Acinar,Pancreas-Alpha,Pancreas-Beta,Pancreas-Delta,Pancreas-Duct,Prostate-Ep,Skeletal-Musc,Small-Int-Ep,Smooth-Musc,Thyroid-Ep
+ filterType.cellTypes multipleListOr
+ filterLabel.cellTypes Filter by cell type
+ filterByRange.cellTypeCount on
+ filter.cellTypeCount 1:22
+ filterLimits.cellTypeCount 1:22
+ filterLabel.cellTypeCount Number of cell types sharing the region